Pyraloidea

Pyraloidea
European corn borer, Ostrinia nubilalis
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Lepidoptera
(unranked):	Obtectomera
Superfamily:	Pyraloidea
Families
Pyralidae Crambidae
Diversity
c.215 genera, roughly 16,000 described species

The Pyraloidea (pyraloid moths) are a moth superfamily containing about 16,000 described species worldwide (Munroe & Solis 1998), and probably at least as many more remain to be described. They are generally fairly small moths.

This superfamily used to contain the Hyblaeidae, Thyrididae, Alucitidae (plus Tineodidae), Pterophoridae, and Pyralidae. Currently, the Crambidae are usually separated from the Pyralidae, but the first four families are now each split off as a distinct superfamily. Some genera (e.g. Hydriris, Micronix and Tanaobela) still defy easy classification and have been variously assigned to the Crambidae or the Pyralidae.

Among all Lepidoptera, pyraloids show the most diverse life history adaptations. The larvae of most species feed on living plants either internally or externally as leaf rollers, leaf webbers leaf miners, borers, root feeders, and seed feeders. Some species live parasitically in ant nests (Wurthiinae), prey on scale insects (certain Phycitinae), or live in the nests of bees (Galleriinae). The larvae of the Acentropinae are adapted to life under water, and certain Phycitinae and Pyralinae are adapted to very dry environments and their larvae feed on stored food products. Others feed on animal detritus such as carrion and feces.

With such a variety of living habits, pyraloids are used in biodiversity studies (Schulze & Fiedler 2003). Some species are of economic importance, e.g.:

• rice stem borers (Chilo spp.; Scirpophaga spp.)
• sod grass webworms (different species of Crambinae)
• Indian meal moth (Plodia interpunctella)
• European corn borer (Ostrinia nubilalis)
• Indo-Australian coconut spike moth (Tirathaba rufivena)
• Cacao moth (Ephestia elutella)
• Mediterranean flour moth (Ephestia kuehniella)
• wax moths (Achroia grisella, Galleria mellonella)
• rice moth (Corcyra cephalonica).

References

Munroe, E. G. & M. A. Solis 1998: The Pyraloidea. Pp. 233–256. – In: Kristensen, N. P., Lepidoptera, Moths and Butterflies. Volume 1: Evolution, systematics, and biogeography. - In: M. Fischer (ed.), Handbook of Zoology. Volume IV Arthropoda: Insecta, Part 35. – Walter de Gruyter, Berlin.

Schulze, C. H. & K. Fiedler 2003: Vertical and temporal diversity of a species rich moth taxon in Borneo. Pp. 69–88. – In: Basset, Y., V. Novotny, S. E. Miller & R. L. Kitching, Arthropods of tropical forests. Spatio-temporal resource use in the canopy. – Cambridge University Press.

External links

• Global Information System on Pyraloidea (GlobIZ)
• Scielo Solis, M. Alma. Phylogenetic studies and modern classification of the Pyraloidea (Lepidoptera ). Rev. Colomb. Entomol., Jan./June 2007, vol.33, no.1, p. 1-8. ISSN 0120-0488. Free download pdf
• Images of Pyraloid Moths in New Zealand